1. Department of Genetics, Evolution, and Environment, University College London , Gower Street , London WC1E 6BT, UK
2. School of Biomedical Engineering, Capital Medical University , Beijing 100069 , China
3. Department of Statistics and Data Science, China Southern University of Science and Technology , Shenzhen, Guangdong 518055 , China
4. Department of Evolution and Ecology, University of California , Davis, CA 95616 , USA