1. Department of Genetics, Evolution, and Environment, University College London, London WC1E 6BT, United Kingdom
2. Department of Statistics and Data Science, China Southern University of Science and Technology, Shenzhen 518055, China
3. Department of Intelligent Medical Engineering, School of Biomedical Engineering, Capital Medical University, Beijing 100069, China
4. Department of Evolution and Ecology, University of California, Davis, CA 95616