1. Centre for Biological Diversity, School of Biology, University of St Andrews , St Andrews , UK
2. Department of Biology, Texas State University , San Marcos, TX , USA
3. The Data Analysis Group, School of Life Sciences, University of Dundee , Dundee , UK
4. School of Medicine, University of North Haugh , St Andrews KY16 9TF , UK
5. Instituto de Ecologia, Universidad Nacional Autónoma de México, Ciudad Universitaria , Mexico City CdMx , Mexico