1. The Key Laboratory of Forest Protection, State Forestry Administration of China, Research Institute of Forest Ecology, Environment and Protection, Chinese Academy of Forestry, Beijing 100091, China
2. Institute of Botany, Beijing Botanical Garden, Beijing 100093, China
3. State Key Laboratory of Tree Genetics and Breeding, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, China
4. Forestry Resources Protection Institute, Jilin Provincial Academy of Forestry Sciences, Changchun 130031, China