1. The David Hide Asthma and Allergy Research Centre, Newport, Isle of Wight, UK
2. Clinical and Experimental Sciences, University of Southampton, Southampton, UK
3. NIHR Biomedical Research Centre, University Hospitals Southampton NHS Foundation Trust, Southampton, UK
4. Human Development and Health, University of Southampton, Southampton, UK
5. Division of Epidemiology, Biostatistics, and Environmental Health, University of Memphis, Memphis, TN, USA
6. Department of Large Animal Clinical Sciences, Michigan State University, East Lansing, MI, USA