1. The New Zealand Institute for Plant and Food Research Limited , Private Bag 92169, Auckland 1142, New Zealand
2. Key Laboratory of Horticultural Plant Biology (Ministry of Education) , College of Horticulture and Forestry, Huazhong Agricultural University, 1 Shizishan Street, Wuhan, 430070, China
3. Zhengzhou Fruit Research Institute , Chinese Academy of Agricultural Sciences, 32 Gangwan Road Zhengzhou 450009, China