1. Department of Biology, Tokyo Gakugei University, Koganei-shi 184-8501, Japan
2. These authors contributed equally to this work.
3. Laboratory of Cell Dynamics, Graduate School of Bioagricultural Sciences, Nagoya University, Nagoya 464-8601, Japan
4. Present address: Department of Biology, Tokyo Gakugei University, Koganei-shi 184-8501, Japan.
5. Present address: Hatoyama High School, Saitama, Hiki-gun, Hatoyama-machi 350-0313, Japan.
6. Department of Biological Sciences, Graduate School of Science, University of Tokyo, Bunkyo-ku, Tokyo 113-0033, Japan
7. Okazaki Institute for Integrative Bioscience, National Institutes of Natural Sciences, Okazaki 444-8787, Japan