1. Graduate School of Life Science, Hokkaido University, Sapporo, 060-0810 Japan
2. These authors contributed equally to this work.
3. Department of Mathematics, Hokkaido University, Sapporo, 060-0810 Japan
4. King Abdullah University of Science and Technology (KAUST), Computational Bioscience Research Center (CBRC), Thuwal 23955-6900, Saudi Arabia
5. RIKEN Center for Sustainable Resource Science, Yokohama, 230-0045 Japan
6. Present address: National Institute of Agrobiological Sciences, NARO, Tsukuba 305-8602 Japan.
7. Graduate School of Information Science, Nara Institute of Science and Technology, Ikoma, 630-0192 Japan
8. Faculty of Science, Hokkaido University, Sapporo, 060-0810 Japan
9. Graduate School of Agriculture, Hokkaido University, Sapporo, 060-8589 Japan