1. Cryobiofrontier Research Center, Faculty of Agriculture, Iwate University, Ueda 3-18-8, Morioka, 020-8550 Japan
2. NARO Tohoku Agricultural Research Center, 4 Akahira, Shimo-Kuriyagawa, Morioka, 020-0198 Japan
3. Department of Plant-Biosciences, Faculty of Agriculture, Iwate University, Ueda 3-18-8, Morioka, 020-8550 Japan