1. Oki Marine Biological Station, Shimane University, Oki, Shimane, Japan
2. Atmosphere and Ocean Research Institute, University of Tokyo, Kashiwa, Chiba, Japan
3. Institute of Biological, Environmental and Rural Science (IBERS), Aberystwyth University, Penglais, Aberystwyth, UK
4. Department of Ichthyology and Fisheries Science, Rhodes University, Grahamstown, South Africa