1. Atmosphere and Ocean Research Institute University of Tokyo Chiba Japan
2. School of Marine Science and Technology Tokai University Shizuoka Japan
3. Department of Life Sciences Shimane University Shimane Japan
4. Department of Ichthyology and Fisheries Science Rhodes University Grahamstown South Africa
5. Institute of Biological, Environmental and Rural Sciences (IBERS) Aberystwyth University Aberystwyth UK