1. Department of Entomology, Center for Pollinator Research, Pennsylvania State University , University Park, PA 16802 , USA
2. U.S. Department of Agriculture, Agricultural Research Service (USDA-ARS), Pollinating Insects Research Unit, Utah State University , Logan, UT 84322 , USA
3. Laboratório de Sistemática de Insetos, Instituto de Ciências Biológicas, Universidade Federal de Minas Gerais , Belo Horizonte, MG , Brazil