1. Graduate School of Biostudies, Kyoto University , Kyoto 606-8501, Japan
2. Department of Biochemistry and Molecular Genetics, University of Colorado School of Medicine , Denver, CO 80045, USA
3. RNA Bioscience Initiative, University of Colorado School of Medicine , Aurora, CO 80045, USA
4. Department of Genetics, University of North Carolina , Chapel Hill, NC 27599, USA
5. Department of Biology, University of North Carolina , Chapel Hill, NC 27599, USA