1. School of Biological Sciences, University of Canterbury, Private Bag 4800, Christchurch 8140, New Zealand
2. Faculty of Science and Technology and Athabasca River Basin Research Institute, Athabasca University, 1 University Dr. Athabasca, Alberta T9S 3A3, Canada
3. Department of Biological Sciences, University of Alberta, 11445 Saskatchewan Dr. Edmonton, Alberta T6G 2E9, Canada