1. Groningen Institute for Evolutionary Life Sciences, University of Groningen , Groningen , The Netherlands
2. Department of Medical Microbiology and Virology, Carl von Ossietzky University Oldenburg , Oldenburg , Germany
3. Institut des Sciences de l’Evolution de Montpellier UMR5554, Université de Montpellier, CNRS-IRD-EPHE-UM , Montpellier , France
4. Groningen Biomolecular Sciences and Biotechnology Institute, University of Groningen , Groningen , The Netherlands