1. Department of Biology, Center for Wildlife Forensics and Conservation Studies, University of Central Oklahoma , 100 North University Drive, Edmond, OK 73034 , USA
2. Division of Ecology and Evolution , , 46 Sullivan’s Creek Way, Acton, ACT 2601 , Australia
3. Research School of Biology, Australian National University , , 46 Sullivan’s Creek Way, Acton, ACT 2601 , Australia