1. Department of Biological Sciences, University of Toronto Scarborough , 1265 Military Trail, Toronto, Ontario , Canada
2. Department of General and Systematic Zoology, University of Greifswald, Loitzer Strasse 26, 17489 Greifswald , Germany
3. PSL Université Paris: EPHE-UPVD-CNRS, USR 3278 CRIOBE, Université de Perpignan , 52 Avenue Paul Alduy, CEDEX 9, 66860 Perpignan , France