1. Doctoral Programme in Wildlife Biology, Faculty of Biological and Environmental Sciences, University of Helsinki , Viikinkaari, Helsinki , Finland
2. Research Programme in Organismal and Evolutionary Biology, Faculty of Biological and Environmental Sciences, University of Helsinki , Viikinkaari, Helsinki , Finland
3. HiLIFE Helsinki Institute of Life Science, University of Helsinki , Viikinkaari, Helsinki , Finland
4. School of Natural Sciences, Macquarie University , 14 Eastern Road, Sydney, NSW 2109 , Australia