1. Department of Neurobiology, Physiology and Behavior, University of California Davis , One Shields Avenue, Davis, CA 95616 , USA
2. Animal Behavior Graduate Group, University of California Davis , One Shields Avenue, Davis, CA 95616 , USA
3. Department of Biological Sciences, North Dakota State University , 1340 Bolley Drive, Fargo, ND 58102 , USA
4. Smithsonian-Mason School of Conservation & Department of Biology, George Mason University , 1500 Remount Rd, Front Royal, VA 22630 , USA