1. Department of Biology, McMaster University , 1280 Main St. W, Hamilton, Ontario L8S 3L8 , Canada
2. School of Earth, Environment & Society, McMaster University , 1280 Main St. W, Hamilton, Ontario L8S 3L8 , Canada
3. Department of Biochemistry, The University of Western Ontario , 1151 Richmond St., London, Ontario N6A 5C1 , Canada