1. Department of Evolutionary Biology and Environmental Studies, University of Zurich-Irchel Winterthurerstrasse 190, Zurich, Switzerland
2. Department of Biology and Institute of Integrative Biology, ETH Zurich, Universitätsstrasse 16, 8092 Zurich, Switzerland
3. Biological and Environmental Sciences, University of Stirling, Stirling, Scotland, UK
4. Biology and Environmental Sciences, University of Gothenburg, Gothenburg, Sweden