1. Centre for Biodiversity Dynamics, Department of Biology, Norwegian University of Science and Technology Høgskoleringen 5 , NO-7491 Trondheim , Norway
2. Department of Natural History, NTNU University Museum, Norwegian University of Science and Technology , Trondheim , Norway
3. Department of Biology, Norwegian University of Science and Technology Høgskoleringen 5 , NO-7491 Trondheim , Norway