1. the Chinese Academy of Sciences CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, , Wuhan 430074, Hubei, China
2. University of Chinese Academy of Sciences College of Life Sciences, , Beijing 100049, China
3. Cornell University School of Integrative Plant Science, Section of Plant Biology and the L.H. Bailey Hortorium, , Ithaca, NY 14853 USA
4. Boyce Thompson Institute BTI Computational Biology Center, , Ithaca, NY 14853, USA