1. Department of Biology, University of York, Wentworth Way, York YO10 5DD, UK.
2. Biodiversity Research Center, Academia Sinica, 128 Academia Road, Section 2, Nankang Taipei 115, Taiwan.
3. School of Biological and Biomedical Sciences, and Institute of Hazard, Risk and Resilience, Durham University, South Road, Durham DH1 3LE, UK.
4. Centre for Ecology & Hydrology, Crowmarsh Gifford, Wallingford, Oxfordshire, OX10 8BB, UK.