1. Department of Integrated Applied Life Science, Integrated Graduate School of Medicine, Engineering, and Agricultural Sciences, University of Yamanashi, Kofu, Yamanashi, Japan
2. Graduate School of Life and Environmental Sciences, University of Yamanashi, Kofu, Yamanashi, Japan
3. Kiwa Laboratory Animals Co., Ltd., Kiminocho, Wakayama, Japan
4. Advanced Biotechnology Center, University of Yamanashi, Kofu, Yamanashi, Japan
5. Faculty of Life and Environmental Sciences, University of Yamanashi, Kofu, Yamanashi, Japan