1. The Sainsbury Laboratory, University of East Anglia, Norwich Research Park , Norwich NR4 7UH, UK
2. Laboratory of Crop Evolution, Graduate School of Agriculture, Kyoto University , Mozume, Muko, Kyoto, 617-0001 Japan
3. JST-PRESTO , 4-1-8, Honcho, Kawaguchi, Saitama, 332-0012 Japan