1. Department of Science Education, Graduate School of Education, Hiroshima University, Higashihiroshima, Japan
2. Graduate School of Human and Environmental Studies, Kyoto University, Kyoto, Japan
3. Kitakyushu Museum of Natural History and Human History, Kitakyushu, Japan
4. Fauna Biodiversity Section, Research, Development & Innovation Division, Forest Department Sarawak, Kuching, Sarawak, Malaysia
5. Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand