1. Key Laboratory of Aquatic Biodiversity and Conservation of Chinese Academy of Sciences, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, China
2. Institute of Evolution and Marine Biodiversity, Ocean University of China, Qingdao 266003, China
3. Department of Zoology, Faculty of Science, Charles University, Prague 128 43, Czech Republic
4. Department of Biology, University of Pisa, Pisa Via Luca Ghini 13, 56126, Italy