1. Pacific Biological Station, Fisheries and Oceans Canada , Nanaimo, BC V9T 6N7 , Canada
2. Bedford Institute of Oceanography, Fisheries and Oceans Canada , Dartmouth, NS B2Y 4A2 , Canada
3. Department of Integrative Biology, University of Guelph , Guelph, ON N1G 2W1 , Canada
4. Institute of Ocean Sciences, Fisheries and Oceans Canada , Sidney, BC V8L 5T5 , Canada
5. Department of Forest and Conservation Sciences, University of British Columbia , Vancouver, BC V6T 1Z4 , Canada