1. Centre for Environment, Fisheries and Aquaculture Science , Pakefield Rd, Lowestoft NR33 0HT , UK
2. School of Earth and Environment, Sustainability Research Institute, University of Leeds , Woodhouse, Leeds LS2 9JT , UK
3. Collaborative Centre for Sustainable Use of the Seas, University of East Anglia , Norwich Research Park, Norwich NR4 7TJ , UK