1. Graduate School of Oceanography, University of Rhode Island, South Ferry Road, Narragansett, RI 02882, USA
2. The Nature Conservancy, South Ferry Road, Narragansett, RI 02882, USA
3. The Bren School, University of California Santa Barbara, CA 93106, USA
4. Marine and Freshwater Research Centre, Galway-Mayo Institute of Technology, Dublin Road, Galway, H91 T8NW, Ireland