1. Key Laboratory of Mariculture, Ministry of Education, Ocean University of China , Qingdao 266003 , China
2. Fisheries and Oceans Canada, Pacific Biological Station , 3190 Hammond Bay Rd, Nanaimo, BC V9T 6N7 , Canada
3. Marine College, Shandong University , Weihai 264209 , China
4. IRD, Univ Montpellier, Ifremer, CNRS, MARBEC , Montpellier 30171, 34203 , France
5. Frontiers Science Center for Deep Ocean Multispheres and Earth System (FDOMES), Ocean University of China , Qingdao 266003 , China
6. Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao National Laboratory for Marine Science and Technology , Qingdao 266071 , China