1. Department of Wildlife, Fish, and Conservation Biology, University of California, Davis, 1 Shields Avenue, Davis, CA 95616, USA
2. National Marine Fisheries Service, Northwest Fisheries Science Center, 2725 Montlake Boulevard E, Seattle, WA 98112, USA
3. Department of Environmental Science and Policy, University of California, Davis, 1 Shields Avenue, Davis, CA 95616, USA