1. Departments of Entomology and Biology, The Pennsylvania State University, State College, Pennsylvania 16802, USA;
2. Department of Ecology, Evolution and Marine Biology, University of California, Santa Barbara, California 93106, USA
3. IFREMER, Centre de Recherche Halieutique Méditerranden et Tropical, Boulevard Jean Monnet, BP 171, 34203 Sète cedex, France