1. Cathy Dichmont Consulting, Bribie Island 4507, Australia
2. CSIRO Oceans and Atmosphere, Hobart, Tasmania 7000, Australia
3. CSIRO Oceans and Atmosphere, Queensland Biosciences Precinct, 306 Carmody Rd, St Lucia, Brisbane, Queensland 4067, Australia
4. Great Barrier Reef Marine Park Authority, PO Box 1379, Townsville, Queensland 4810, Australia
5. Department of Agriculture and Fisheries, Fisheries Queensland, GPO Box 46, Brisbane, Queensland 4001, Australia
6. Department of Applied Mathematics and Statistics, University of California, Santa Cruz, California, CA 95060, USA
7. Theoretical Ecology Group, University of Bergen, Bergen 9020, Norway