1. Plant Breeding, Center of Life and Food Sciences Weihenstephan, Technische Universität München, 85354 Freising, Germany, 2 Department of Plant Genome and Systems Biology, Helmholtz Center Munich, German Research Center for Environmental Health, 85764 Neuherberg, Germany and 3 College of Science, King Saud University, Riyadh 11451, Kingdom of Saudi Arabia