1. School of Earth Sciences and Engineering, Guangdong Provincial Key Lab of Geological Processes and Mineral Resources, Sun Yat-sen University , China
2. School of Life Sciences, Key Laboratory of Conservation and Application in Biodiversity of South China, Guangzhou University , China
3. School of Ecology, Sun Yat-sen University , China
4. College of Life Sciences, Capital Normal University , China
5. Department of Paleobiology, National Museum of Natural History, Smithsonian Institution , USA
6. Department of Entomology, University of Maryland , College Park , USA