1. Institute of Vertebrate Paleontology, College of Life Science and Technology, Gansu Agricultural University, Lanzhou 730070, China
2. Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum National d’Histoire Naturelle, CNRS, SU, EPHE-PSL, U A, CP50, 45 Rue Buffon, F-75005 Paris, France
3. Institut des Sciences de l’Évolution de Montpellier (UMR 5554), Université de Montpellier, CNRS, F-34095 Montpellier, France
4. Univ. Rennes, CNRS, Géosciences Rennes, UMR 6118, F-35000 Rennes, France
5. College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China
6. Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013, USA