1. Department of Molecular Physiology, Faculty of Life Sciences, Kumamoto University, Honjo 1-1-1, Chuo-ku, Kumamoto-shi, Kumamoto 860-8556, Japan
2. Department of Modomics Biology and Medicine, Institute of Development, Aging and Cancer, Tohoku University, Seriyo-machi 4-1, Aoba-ku, Sendai-shi, Miyagi 980-8575, Japan