1. Department of Biochemistry, McGill University , Montreal , Quebec , Canada
2. The Rosalind & Morris Goodman Cancer Institute, McGill University , Montreal , Quebec , Canada
3. Department of Human Genetics, McGill University , Montreal , Quebec , Canada
4. Institute for the Advanced Study of Human Biology (WPI-ASHBi) , Kyoto , Japan
5. Canadian Center for Computational Genomics,McGill University , Montreal, Quebec, Canada