1. Centre for Discovery Brain Science, Deanery of Biomedical Sciences, University of Edinburgh , Edinburgh EH8 9XD , UK
2. School of Life Sciences, University of Nottingham Medical School , Nottingham NG7 2UH , UK
3. Experimental Medicine and Immunotherapeutics, University of Cambridge , Cambridge CB2 0QQ , UK
4. Spedding Research Solutions SAS , Le Vésinet 78110 , France