1. Department of Biological Sciences, Korea Advanced Institute of Science and Technology , Daejeon 34141 , Republic of Korea
2. Division of Life Sciences, Korea University , Seoul 02841 , Republic of Korea
3. Department of Microbial and Cellular Sciences, University of Surrey , Guildford GU2 7HX, UK
4. School of Biological Sciences, Seoul National University , Seoul 08826 , Republic of Korea