1. School of Life Sciences and Allied Health Professions, Kamuzu University of Health Sciences Blantyre , Blantyre, 265 , Malawi
2. Institute of Infection, Veterinary & Ecological Sciences, University of Liverpool , Liverpool, L69 7ZB , United Kingdom
3. Malawi–Liverpool–Wellcome Programme , Blantyre, 3 , Malawi
4. Department of Microbiology, Moyne Institute of Preventive Medicine, School of Genetics and Microbiology, Trinity College Dublin , Dublin, D02 PN40 , Ireland
5. Department of Public Health and Epidemiology, College of Medicine and Health Sciences, Khalifa University , Abu Dhabi, P.O. Box 127788 , United Arab Emirates
6. Smurfit Institute of Genetics, School of Genetics and Microbiology, Trinity College Dublin , Dublin, D02 PN40 , Ireland
7. Research Department of Infection, Division of Infection & Immunity, University College London , London, WC1E 6BT , United Kingdom