1. Department of Ecology, Evolution and Organismal Biology, Iowa State University , 251 Bessey Hall, 2200 Osborne Drive, Ames, IA 50011 , USA
2. Institut de Biologie de l’ENS (IBENS), École normale supérieure, CNRS, INSERM, Université PSL , 46 rue d’Ulm, 75005 Paris , France
3. Department of Invertebrate Zoology and Parasitology, University of Gdańsk , Wita Stwosza 59, 80-308 Gdańsk , Poland
4. Museum of Nature Hamburg, Leibniz Institute for the Analysis of Biodiversity Change , Martin-Luther-King Platz 3, 20146 Hamburg , Germany