1. College of Life Sciences, Hunan Normal University, Changsha, Hunan 410081, China
2. State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution, School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China
3. School of Life and Environmental Sciences, University of Sydney, Sydney, NSW, Australia
4. Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543, Singapore
5. Department of Biomedical Science and Environmental Biology, Kaohsiung Medical University, Kaohsiung City 80708, Taiwan
6. Department of Organisms and Ecosystems Research, National Institute of Biology, Ljubljana, Slovenia
7. Jovan Hadži Institute of Biology, Scientific Research Centre of the Slovenian Academy of Sciences and Arts, Ljubljana, Slovenia
8. Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA
9. University of Ljubljana, Ljubljana, Slovenia
10. Department of Zoology, National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba-shi, Ibaraki-ken 305-0005, Japan
11. Center of Excellence in Entomology and Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand
12. Department of Zoology, University of Yangon, Kamayut Township, Pyay Road, Yangon 11041, Myanmar
13. Department of Biology, Taungoo Education College, Taungoo 08101, Myanmar
14. Department of Experimental Taxonomy and Genetic Diversity, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, Hanoi, Vietnam
15. Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603 Kuala Lumpur, Malaysia, Xin Xu and Yong-Chao Su contributed equally to this work