1. Centre for Macroevolution and Macroecology, Research School of Biology, Australian National University, Canberra, ACT 0200, Australia
2. School of Biological Sciences, University of Sydney, Sydney, NSW 2006, Australia
3. School of Biological Sciences, Royal Holloway University of London, Egham, Surrey TW20 0EX, UK
4. Sackler Institute for Comparative Genomics, American Museum of Natural History, New York, NY 10024, USA
5. Department of Biology, The Pennsylvania State University, University Park, PA 16802, USA