1. Museum of Zoology and Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, USA
2. Department of Biology, California State University—Dominguez Hills, Carson, CA 90747, USA
3. College of Science & Engineering, James Cook University, Townsville, Queensland 4811, Australia
4. Biodiversity Program, Queensland Museum, South Brisbane, Queensland 4101, Australia
5. Division of Ecology and Evolution, Research School of Biology and Centre for Biodiversity Analysis, Australian National University, Acton, ACT 2601, Australia