1. Center for Computational and Evolutionary Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China
2. State Key Laboratory of Integrated Management of Pest Insects and Rodents, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China
3. Graduate University of Chinese Academy of Sciences, Beijing 100049, China
4. School of Mathematical Sciences, Peking University, Beijing 100871, China
5. Department of Biology, University College London, Darwin Building, Gower Street, London WC1E 6BT, UK