1. Laboratory of Systematic & Evolutionary Botany and Biodiversity, College of Life Sciences Zhejiang University Hangzhou 310058 Zhejiang China
2. Plant Quarantine Station of Lin'an District Hangzhou 311300 Zhejiang China
3. Nanjing Institute of Geology and Palaeontology Chinese Academy of Sciences Nanjing 210008 Jiangsu China
4. Nature Fieldwork Center Okayama University of Science Okayama 700‐0005 Japan
5. Museum of Nature and Human Activities Hyogo 6 chome, Yayoigaoka Sanda Hyogo 669‐1546 Japan
6. College of Life and Environmental Science Wenzhou University Wenzhou 325035 China
7. Department of Botany University of Wisconsin Madison WI 53706 USA