1. Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA; E-mail: lliu@oeb.harvard.edu, sedwards@fas.harvard.edu
2. Department of Biostatistics, Georgia Southern University, Statesboro, GA 30460, USA; E-mail: lyu@georgiasouthern.edu
3. Department of Statistics,The Ohio State University, Columbus, OH 43210, USA; E-mail: dkp@stat.osu.edu